I'm a dumbass.
I went through all your suggestions and found nothing wrong. I decided to go back and look for issues with the board. I didn't think it was the boards, because the issue was symmetrical. So I figured that the issue might be somewhere in the middle of the amp. However, because I built the C4s boards along sides one another, when I missed the step where I connect breg on B and bB, I missed it for both boards.
I added the wire, and now everything is working.
IA on both boards now 515v.
IO on the A board is 297v. IO on the D board is 298v.
32u and 46u both now adjustable to 175v.
Thanks again for your help and infinite patience... Can't wait to knock this out and start on BeePre next when that comes in!