BadDog,
You have mentioned the tube socket LED several times, and that is a good thing. This indicates that when the C4S board hasn't come up to voltage, "turned on," that the tube end of things is just fine. Of course, the problem is in the board.
You are also in luck, you have one good board and one bad board. You can compare both of them with that big magnifying glass you have. Compare the values of the resistors in each position. Your swapping the boards and the problem swapping sides says that the problem is on the board, not the connections. I suggest not beating your head against the wall with that anymore.
Since you have checked the LED orientation several, ok, 10 times let's leave that alone too. You had a couple of transistors swapped. You need to find out from the Bottlehead crew if that might have caused a problem with either of the transistors.