Terminals 4 and 17 are both supposed to be pulled to ground by the 249K ohm resistor connecting them to 5 and 16 respectively. So a bad connection at either end of each resistor could interrupt that, but surprising that both would be affected the same way.
Now if the 2A3 tubes were both plugged in rotated in the same wrong way, you might have built voltage dividers across the tube heater filament, which could explain it. Check to make sure the 2A3's are plugged in with the correct orientation.