Alright, so you have 1.25V at pin 9, this is drawn across a 1.27K resistor, which means that 1mA of current is flowing where there should be 2mA.
1.2V of bias on the tube and 300V at the plate would indicate 40mA of current drawn based on the curves for the 6DJ7.
Either you have shorted transistors in the C4S, or you have lost the ground reference for the grid on that 6FJ7. The grid for section 1 is pin 10 on the 12 pin socket, in is connected to pin 8 by a 220 ohm resistor, then is wired to the volume pot. If any of those connections are loose or poorly soldered, you will lose your ground reference and your voltages will go whacky. This can include all of the connections at the pot itself, so a broken ground wire up by the pot could cause this issue.
If you have shorted transistors on the C4S, this could be confirmed by setting your meter to measure continuity (it will beep when you touch the probes together), then check each pair of legs on each transistor on the C4S board to look for shorts. If you find shorted transistors, then the immediate concern would be why they shorted, which would go back to a poor solder joint.