If rotating the center board also rotated the problem, then there's a trimmer pot, a 2N2222 transistor, and a resistor that are being swapped in that process, as well as the high voltage regulator components. Before swapping the center board again, what voltages do you see at OA and OB on that center board?
I have to be absolutely insistent that you not say anything about lit or dim LEDs moving forward. It's absolutely 100% necessary to have voltage measurements. Dark LEDs don't really tell me anything, while voltage measurements tell me a lot.