Hey Scott,
I'm no expert, but I had the same problem when I first built my BeePre - I had miswired the A B C D terminals on the selector switch. Have you tried all three pairs of inputs on the BeePre? If that doesn't make a difference, I'd double check all the red and black wiring at the input selector switch and the balance and volume pots. I'd also test whether signal is getting to the grid of the right 300B. With amp unplugged and cold, I'd connect one test probe to A3, set the input selector switch to input 1, set the meter for continuity, and connect/touch the other probe to the center pin of red/right rca input 1. If the meter doesn't beep, I'd try to figure out where the break in connection is -- e.g., leaving one probe connected to the rca center pin, remove the other probe from A3 and put it on D (red) of the selector switch, and then on each of the lugs on the balance and volume pots to which a red wire connects.
If the red signal wiring is all good, I repeat the same continuity test for the signal ground, starting with one probe on terminal 8 and the other on the ground/-ve tab of red rca input 1.
Don;t know wither the above will help, but it's worth a shot.
cheers, Derek