No, I never took out the 12AU7. I removed the 7308 and the EF86s, then put the 7308 back in.
Now I have all 4 tubes installed. I think I found the cause of the smoke (a tiny solder whisker between legs of a transistor). Fixed that and now I have:
IA 218 IB 280
OA 163 OB 230
OC 98 OD 53
OKA 99 OKB 59
OKC 2 OKD -.004