I asked Hamad at Amplifier_surgery (eBay) about C1, and he said this:
Dear tomp961,
Hi Tom, C1 may have been an attempt to eat DC spikes or produce a warmer sound. But it will effect sound if a poor capacitor selection is placed there. Any high quality polypropylene (low leakage) cap would work well there. Regards, Hamad
No wonder Paul J was puzzled. Once I have all tubes matching in the two Quad II amps, I'll listen for a difference between Obbligato Gold Premium and Bottlehead F104K. I don't expect to hear a difference, but with the two amps set up for the test, I'll try it.