In direct coupled designs the driver plate is directly connected to the power tube grid, so that voltage affects the operating point of both tubes. The driver usually (at least in my designs) has plenty of headroom and can tolerate a range of plate voltages, but the power tube can't if you want to get the best out of it.
For this reason, the current (v1.1) Paramount incorporates the "soft-start" upgrade board, which has and adjustable driver cathode voltage. The optimum adjustment is dependent on your power line voltage so you have to calculate it, and then you must check it and re-balance if the tube drifts too far - IHMO, +/-5% or about +/-10v. In the present design that means taking the amp to the test area, turning it over, connecting a meter, and adjusting a trimpot on the PC board. Inconvenient, but as I said this was not the primary design application. I guess, like reel-to-reel tape, this is a design for geeks!
To my ears, a cap coupled amp with a really excellent coupling capacitor, and operated within its power capabilities, is audibly comparable to direct coupling. These days I am more enamored of a fully shunt regulated power supply for that last bit of nuance. There is no such current product but I AM working on it!