It's also important to consider the AC component, especially with those parallel feed capacitors. In the Stereomour, for example, we have 300V DC plate to cathode across the cap, but when you put signal through that 2A3, there's potentially 120V of AC plate signal riding on top of that. A 400V-500V rated cap is not going to like this, but the 630V cap has the margin tolerate this abuse.