My best hope was that the 2A3 was drawing excessive current; apparently that is not the problem.
If you haven't, repeat the resistance checks in the hope that is will reveal something. Stop there and post if you find a problem. If not, do this:
At the bottom of page 18, there is a picture of the PSU board, showing the high voltage output of the power supply - the twisted pair of black and red wires. Disconnect the red wire from the board temporarily - this will isolate the rest of the circuit from the power supply, so we can determine whether the problem is on the PSU board or in the rest of the circuit. Without the 2A3s installed, fire it up to see if the fuse blows. If it doesn't, repeat with 2A3s installed. If it still does not blow, do the voltage checks.
If the fuse still blows, get some 1.5 amp fuses - sometimes a different manufacturer will affect the fuse performance.
(I'm doing my best here to avoid messing with the PSU board, which is very difficult to remove from the power transformer without damage.)