That's good information. When you flipped the board around and the low voltage changed from OB to OA, that tells us that the small PC board is working.
As to why the issue is occurring in the first place, some potential causes are debris in or around the tube socket causing a short between pin A6 and A7 (or A6 and A5), what DC resistance do you get from terminal 5 to ground?
Also, to reiterate again, this is a problem that has nothing to do with the LED that isn't lighting. If the LED was shorted out, terminal 5 would read something like 50V, and if the LED was open it would read around 170V.