Well, this tells us that the C4S bits of the PC board are OK (the center bits with the MJE5731A/2N2907 and LED's).
When you have the OA/OB bits jumpered to the outsides of the board, your voltages are sinking like crazy. You've checked the placement of the individual parts, but I'd wonder if there's an issue with the way the board is wired to the tube socket. If the OA and OB pads aren't connected to D1 and D6, but rather to D3 and D8, I could see you getting this problem.
-PB