If you don't see it at the output of the volume pot, then it's likely a bit of imbalance from one of the tubes. Unfortunately, your meter doesn't resolve AC voltage well enough to really tell if this is happening before or after the tubes. I would suggest grabbing a meter with a 2V AC scale on it (let me know where you're located and I can suggest some retailers), then you can resolve your signal levels a bit better and know exactly where things are going awry.
The reason I would suggest this is that 0.28V and 0.32V on the 200V AC scale will both probably show up as 0.3V, but they are 1dB apart.