And just to be absolutely positive, if you take the tube that's out of the amp now and swap it with the tube that's still in the amp, does the low power supply voltage follow that particular tube?
What we learned by the voltage popping up when you removed the plate choke wire is that there's something connected on or around where the blue plate choke wire lands that shouldn't be there, or something is touching that shouldn't be, and that's dragging down the whole power supply and heating up the power supply resistor.