Of course, since swapping the 6N1P's back again didn't change anything again, it was just wishful thinking on my part for me to believe that the problem had resolved itself.
It worked for a little while, then while I was listening, there was a pop in the affected channel, then it went back to the same low-vol distorted output as before.
So, setting up to get the measurements you suggested, I connected the input and multimeter to the common ground at terminal 16. I then set the dmm to DC. Turned the amp on with tubes installed. Only one led lights on the C4s. The instant I touch the "I" (input?) with my meter lead the other LED immediately lights and stays lit.
I turn off the amp. Figuring out that the G on the C4S is likely ground, I attach the ground of the multimeter to the lead labeled "G" on the C4S. I turn the amp back on. Both LEDs are still lit. I take these measurements:
I 302v
O 210v
I am in between locations so I am testing the amp away from its system but I am assuming that everything works properly as long as the LEDs are lit, which they currently are, but won't know for sure until I put it back in my system early next week.
Any ideas for a root cause?