Thank you both so much for helping me troubleshoot.
I didn't see any obvious dirt on any of the pins, but still cleaned everything with some deox-it. Sprayed into the connectors as well. On retry, the same issue was present with the same tube only on the left side. So, no change.
I had some other 12AT7s, so put those in place. I checked the voltages and now the low KREG B voltage is back - 2.85 with the new 12AT7s, unchanged if I switch the tubes around too. I connected some speakers and alas, the crackling sound is back in the left channel when I move the left tube!
Im interpreting all this to mean there has got to be a loose connection somewhere on the left channel. I quadruple checked all the pin connections and they are SO solid but maybe I should just reflow everything.
I will keep hunting and reflowing until it works, unless you guys have any other ideas. Thank you so much for being so supportive!