It seems like just about anything with current running through it can be microphonic. Cold solder joints, bad tube sockets and wires. If fact, one thing I've learned from troubleshooting gear is to tap on each component and wire to track down problems. When I finish a project and even if there aren't any problems, I take a wooden chopstick, turn up the volume and tap on everything to head off trouble. I almost always find something that needs a bit more solder or just needs to be shifted a little.
So, sometimes when I find a tube position that's microphonic with more than one tube, I start by cleaning the socket contacts, making sure that all the pin receivers are tight and check that solder joints are good and don't have any peaks which might tend to arc to another contact (tube pins aren't all exactly the same size so some might move the pins inside of the chassis more than others).