There is a 330K C4S bias resistor on the board, and the 3.3K is in series with it, one end grounded. So if there is 450 volts at the power supply, then there will be 4.5 volts across the 3.3K resistor. Running 1.5mA the power rating should be larger than 0.00675 watts. :^)
The 300K is grounded, and goes from the driver plate to ground. Sticking a 3K in the ground end gives 2 volts if the driver plate is at 200v. Again, no power issues. The Zener is just there to make sure that whatever connection goes outside the chassis, stays at a low voltage - yes, cathode to positive, anode to ground.
The bias adjustment on the driver occurs in the cathode circuit, where voltages are normally no more than 4 volts. The 431 will reject noise on that line, but it IS in the cathode, a high-gain position, so I suggested the shielded wire out of caution.
Get Paully to bring the manual; a glance at the circuit should make this more clear.