OK, sorry for this:
1) In the original design, that resistor should not be there; it is an error in the manual.
2) With the "soft-start" 5670 driver upgrade (and Paramount v1.1), that resistor is back, but is now 300K, is shown on the driver schematic, and is installed prior to the coupling cap not after. (It is actually installed as a jumper on the PC board.)
3) The circuit error is still there in the v1.1 manual, unfortunately - it still shows the 249K resistor for the 2A3, and it should still not be there.
I do recommend the new driver and board for any 2A3 or 45 Paramount, for two reasons:
1) It protects the output tube grid from transient high voltage at startup; this has caused arcing in a (very few) tubes. As far as I know, this is only a severe problem with the EML 2A3 (and I expect, 45) tubes which should NEVER be used without this change.
2) It allows the precise driver plate voltage to be set to correspond to your house's power line voltage, optimizing the operating point. The operating point of direct coupled circuits is usually very sensitive to the power line voltage.
If you have the original design and you use an old-stock 45 or 2A3, or the 2A3 we supply, you are not likely to have any arcing problem. If you have the original design and your power line is close to 120 volts and your 12AT7 is close enough to the spec, then you will probably be about right on operating point. You can measure the driver and output tube plate voltages and post them here; I'll check them against my calculations and if they are not close to optimal provide suggestions for improving them. This would usually be just adding a small resistor in series with the LED on the driver cathode.