I can't explain the arc the first time. If the speakers are silent for the first 5-10 seconds now, the soft start must be resetting properly. It "resets" when the 10000uF cap on the board discharges through the 33K resistor adjacent to it, so there is no mechanical reset function.
If it ever arc or pops again, immediately replace the 45s with the stock 2A3s and don't put them back until you have identified and solved the problem.