These voltage measurements look to me like they indicate a few problems still. I'll describe what I see from the problem measurements. This post is long because I wanted to describe my reasoning.
1) 269.1 vac
2) 269.4 vac
These are the power transformer secondary, and should be 330v each. If this is accurate and the transformer is working correctly, it indicates your power line voltage is a bit less than 100 volts, where it should be 120 volts. Are your lights very dim? :^) Seriously, I would make sure about this first, it seems on the face of it unlikely but could explain all the problems. Check the power line voltage, and then the filament voltage (AC) across each tube - between A1-A4, and between B9-B4/5. And if you haven't noticed a 20% low power line voltage already with other appliances, then make sure your meter is working properly!
5) 330 vdc
15) 316.9 vdc
A1) 51.2 vdc
A2) 306.5 vdc
All the above voltages are 20% low, consistent with a 20% low line voltage.
A3) 125 vdc (have an auto ranging DMM triple checked and it is reading vac not mvac)
The A1 and A4 voltages indicate the 2A3 is conducting properly, so this voltage cannot be 125vDC - the DC component must be quite small. If it is really reading 125vAC then it may be oscillating badly. (In that case, it's also possible that the meter actually measured peak voltage on teh DC scale, and would read a DC voltage when there is no such voltage on an average basis. This is unlikely but not unheard of.) Make sure the corrected grid stopper wiring has been installed - there should be a 270K resistor from T17 to ground at T14 (NOT A3 to ground as shown in the original manual) and a 499 ohm resistor from T17 to A3. Nothing else should be connected to A3.
It is conceivable that if this is oscillating badly, it could bring the whole power supply voltage down, and the 2A3 would be getting quite hot.
A4) 50.9 vdc
B6) 246 vdc
B8) 2.08
Left most term) 226.4 vdc
The "left most terminal" cannot have a lower voltage than B6, so the voltage must have drifted while you were measuring. It may indicate your power line voltage is fluctuating as well as being low, or it could indicate destructive oscillations that are heating or otherwise causing components to drift.
The fact that this voltage is about the same as the driver plate (B6) indicates the C4S is not able to force its target current through the tube so there is no compliance for positive peaks at the driver output. This would cause severe distortion and low output, it would also couple power supply ripple to the 2A3 grid and cause a good deal of hum.
I mentioned a few possibilities in my previous post, but here are two new ones: A) the left most terminal should be at 295v, giving it a much greater opportunity to exceed B6 by the minimum of 60-70 volts, and B) if the heater voltage is also 20% low the 12AT7 cathode may not be hot enough to provide the necessary current.
Sorry for the long post and many options. I hope it gives you some things to check that will lead to a solution.