Yeah, your 6C45's still aren't drawing any current, but at least you have them heated!
This issue points to an error on the center regulator board, in the trim pot/2N2222A wiring. I don't have one of those PC boards in front of me to be 1000% sure that your accidental miswire of the trim pots will have no effect.
How about the OB/OA voltages on the center PC board?