That's so very odd.
The twisted pair of red/black wires that leave power transformer terminals 4 and 5 are what make the tubes glow, and they stop off at B7/B8 first, then go up to A4,5/A9.
If the 6080 isn't lighting, maybe there's a loose wire on the octal socket at B7/B8. It's also possible that a pin isn't making good contact.
In either case, there's no reason to focus on the Speedball boards when troubleshooting this issue for the moment.
-PB