Sorry, it's been about a month since I posted that, and I don't see any errors. There is something pretty wonky going on here, you have 12V at the cathode of the offending side, the correct resistor in place, and 440V on the plate. This doesn't jive with the curves for a 12AT7.
This leaves us at flaky solder joints, which is very likely your issue. You can also swap the connections at OA and OB, and if the offending voltage stays put, then you know the C4S board is working.
-PB