OK, let's take a small sideways step. Carefully check the AC volts across power transformer terminals 4 and 5, which supply the heater current. Should be around 6.3VAC. Just want to verify the meter calibration.
If that is OK, I think maybe it is time to consider replacing the PN2907 that was in backwards. I don't have an exact scenario in my head of how one 2907 would throw both sides off and pull the cathode LEDS up from 1.56V, but that should more or less take you back to where things went haywire.
Another thing you can try is pulling the 6080 and remeasuring DC voltages OA, OB and A3 and A8. I don't imagine they will change, but it might give some more info.