The next step would be to just remove the large PC board, then run the amp again with just the 12AU7 and the small board.
With the big board out, you can set your meter to DC resistance and measure between each pair of legs on each TIP50 and each 2N2222 transistor. You will likely find that some pairs of legs are shorted together, indicating that a transistor needs to be replaced.
Since the TIP50 transistors were properly mounted and working at some point, it's totally possible that both the TIP50 and 2N2222 where you had your slip are now shorted (thus providing a ground reference on the collector of the TIP50).
With the big board out, we can focus on restoring proper operation to the small PCB.
-PB