You'll want to check a whole bunch of things on your build. First of all, you can measure the R1 resistors with them installed on the board. Are they 237 ohms?
Triple check all of the LM431 and PN2907 transistors to be sure they are in the correct spots.
Since the LEDs are on, the board is passing current, and the high voltage makes it look like a lot of current is being passed. This suggests an issue with the center section of that middle C4S board. Since the LEDs are on and you have appropriate grid to ground resistance on pins 2 and 7 on that driver tube socket, that just leaves the C4S board as the issue.
It would certainly be worth taking the time to go over all the build steps in the manual involving constructing that board and installing it.