The low voltage across the LEDs means not a lot of current is being drawn across it, but you have the correct current being dropped across the 22K resistor on that side of the amp. This could be caused by a flaky solder joint at A4/A5 making half the tube not heat up properly, or possibly a different loose solder joint in the amp, or even the lugs on the pot where the black wires connect may not be well soldered.
You're close to having a working amp, really you should just reflow all the solder joints in the amp other than the power switch and LEDs just to be certain they are all solid.