The ground lugs on the pot are connected with a piece of wire to terminal 12. Since you have 180V between those two points, there is a discontinuity in your black ground wire that is causing your amp to not work properly. As I suggested before, it's extremely common to find this exact issue where the two black wires connect to the headphone jack, and commonly one will not be captured by solder.
From what you are showing, we can be 10000% sure that you have a loose/poorly soldered black wire in your stock build that has nothing to do with the Speedball installation (other than likely being disturbed in the process of fitting that first PC board).
There are only a small handful of solder joints on the ground bus, why not reheat them all? That would be terminal 3, the junction of the black wires on the HP jack, terminal 12, terminal 14, and terminal 20.