Alright, so the center shunt reg board is grounded, receiving power, and there is some voltage present at the 12AU7 cathode.
Now we will be interested in the outboard boards, as we really, really want them to draw some current so the center board can work properly.
First of all, we want to see that the outboard 12AU7's are glowing (seems obvious, but just good to check).
Next, a red wire runs to the "A" side on each of the outer green PC boards to the "I" pad. Measure the voltage here, it should be your ~180V.
A black wire runs to the "B" side on each of the outer green PC boards to the "I" pad. Measure the voltage here, it should be 0. Measure the resistance between these pads and ground, it should be nearly 0. (0.3 in your case)
This would confirm that you have voltage coming into that board. Next, I would want to ensure that the "G" to "G" jumpers are present.
Lastly, if everything above checks out, you can be very suspicious about the LED orientation.
-PB