Hi David,
Sorry to hear my comment above didn't lead anywhere helpful. I had a further thought, borrowing from PB's suggestions in my thread. Have you tried firing up the amp without any 12au7 in the b-socket?
I was slow to take up this suggestion (simply forgot about it) when debugging my own unit, and it turned out to be pivotal since I was focusing on the C4S board when my issue was really on the regulator board.
As I understand it (and I may well be oversimplifying), pulling the 12au7 basically takes the regulator board out of the picture and helps isolate your issue. If your C4S boards still measure differently, then they clearly become the focus. Alternatively, if both C4S boards suddenly perform the same (as occurred in my case), then you can focus on the regulator board.