The 431 regulators are the most sensitive parts on the boards, and you could certainly try replacing them. Since the amp has at least correct DC voltages with no 6AQ5s, that tends to point to the 431 regulators being problematic.
Still, the amp should work properly (perhaps with a little additional noise) with no 6AQ5s installed, and since yours doesn't that would suggest that there are additional issues beyond the high voltage regulator. With the replaced sockets, will the amp work properly with no 6AQ5s? (Be sure to confirm that the voltages are OK first, with OA on the big boards and IA/IB on the smaller C4S board being more like 200V rather than 150V)